The RSN element in messages 2 and 3, plus the wrapped group key in message 3. Empty in messages 1 and 4.
Message 2 echoes back the RSN element the station sent in its association request, and message 3 echoes the one from the beacon. That is the downgrade protection: if an attacker altered either element in the unprotected frames earlier, the echo will not match what was seen and the handshake fails.
EAPOL-Key > Key Data